Actors in an earlier role you just didn't recognise

Classic ignorance from me that it took me ages to recognise that the glamorous Donna Reed who played the love interest in Its A Wonderful Life was in fact the second Miss Ellie in Dallas.

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I watched an episode of open all hours on Monday (I think it was...hubby watches it every weekday) and the bloke who played Boycie from Only fools and Horses was a delivery driver
 
Andrew Scott as a GI in Band of Brothers
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Only spotted him when I watched it a second time, he's only in one episode before stepping on a landmine (I guess first time around he wasn't well known?)
 
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